This song is about a man who goes to a strip club to help him get over a girl. While there, he realizes that this isn't what he wants, and what he really wants is the girl: "Well, I may have faked it, and I wouldn't be caught dead in this place." He recognizes that the girl is playing hard-to-get ("Isn't this exactly where you want me... praying for love in a lapdance") and doesn't like it.
The title comes from the movie Closer, where Natalie Portman's character says, "Lying is the most fun a girl can have without taking her clothes off, but it's better if you do." This line also inspired the title of Panic At The Disco's song, "Lying Is The Most Fun A Girl Can Have Without Taking Her Clothes Off."
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